| MACHINE GUN: DAMAGE | +10 % damage per level. | 5 levels |
| SONIC SHRAPNEL: DAMAGE | +10 % damage per level. | 5 levels |
| TESLA: DAMAGE | +10 % damage per level. | 5 levels |
| MACHINE GUN: MAGAZINE | +1 bullet in the magazine per level. | 5 levels |
| MACHINE GUN: FIRE RATE | -6 % time between shots per level. | 5 levels |
| MACHINE GUN: RELOAD | -10 % reload time per level. | 5 levels |
| MACHINE GUN: CRITICAL | +2 % critical chance per level. | 5 levels |
| TESLA: MAGAZINE | +1 beam in the magazine per level. | 5 levels |
| TESLA: +1 TARGET | The Tesla's beam chains 1 more target (max. 5). | 3 levels |
| TESLA: FIRE RATE | -6 % time between beams per level. | 5 levels |
| TESLA: RELOAD | -10 % reload time per level. | 5 levels |
| TESLA: JUMP CHANCE | More chance that the Tesla's beam jumps. | 5 levels |
| TESLA: RANGE | +5 % range per level. | 5 levels |
| TESLA: JUMP DAMAGE | Chained targets take +10 % more damage per level. | 5 levels |
| TESLA: SLOW | The beam slows by 10 % per level for 2 s. | 5 levels |
| MACHINE GUN: RANGE | +5 % range per level. | 5 levels |
| SONIC SHRAPNEL: RANGE | +5 % range per level. | 5 levels |
| SONIC SHRAPNEL: FIRE RATE | -6 % time between bursts per level. | 5 levels |
| SONIC SHRAPNEL: PELLET COUNT | +1 pellet per hole per level. | 3 levels |
| SONIC SHRAPNEL: RELOAD | -10 % reload time per level. | 5 levels |
| MACHINE GUN: HEALTH | +1 health (1 more hit) per level. | 5 levels |
| SONIC SHRAPNEL: HEALTH | +1 health (1 more hit) per level. | 5 levels |
| TESLA: HEALTH | +1 health (1 more hit) per level. | 5 levels |
| MACHINE GUN: TURN SPEED | +10 % turn speed per level (also the mech). | 5 levels |
| SNIPER: DAMAGE | +10 % damage per level. | 5 levels |
| SNIPER: MAGAZINE | +1 bullet in the magazine per level. | 5 levels |
| SNIPER: FIRE RATE | -6 % time between shots per level. | 5 levels |
| SNIPER: RELOAD | -10 % reload time per level. | 5 levels |
| SNIPER: CRITICAL | +2 % critical chance per level. | 5 levels |
| SNIPER: RANGE | +5 % range per level. | 5 levels |
| SNIPER: HEALTH | +1 health (1 more hit) per level. | 5 levels |
| SNIPER: TURN SPEED | +10 % turn speed per level (also the mech). | 5 levels |
| ORBITAL SUPPORT: DAMAGE | +10 % bombardment damage per level. | 5 levels |
| ORBITAL SUPPORT: RELOAD | -10 % time between bombardments per level. | 5 levels |
| ORBITAL SUPPORT: AREA | +5 % explosion radius per level. | 5 levels |
| ORBITAL SUPPORT: BEAM COUNT | +1 beam per bombardment, each in a different area. | 1 levels |
| ORBITAL SUPPORT: HEALTH | +1 health (1 more hit) per level. | 5 levels |
| ABDUCTOR: CAPTURE SPEED | -6 % capture laser time per level. | 5 levels |
| ABDUCTOR: FARM CAPACITY | +1 maximum animal on the farm per level (from 2 up to 10). | 8 levels |
| ABDUCTOR: HEALTH | +1 health (1 more hit) per level. | 5 levels |
| ABDUCTOR: RANGE | +5 % range per level. | 5 levels |
| ABDUCTOR: PREY SIZE | Level 1: captures medium animals. Level 2: also large ones (without this upgrade, only small). Requirement: HEALTH, CAPACITY and RANGE complete. | 2 levels |
| ABDUCTOR: EXPAND FARM | Expands the farm's land and capacity: +5 animals per level. Level 4: double the area and a second trough. | 4 levels |
| ABDUCTOR: CHOOSE TARGET | Unlocks the TARGET dropdown in the tower menu: choose which animal it captures. Without it, it attacks any. | 1 levels |
| ATOMIC BOMB: AMOUNT | With this node (and its 4 requirements maxed: the machine gun's TRANSFORMATION, the Tesla's SLOW, the sniper's PILOTING and the orbital support's BEAM COUNT) the atomic bomb in slot 9 is unlocked: 1 per battle, +1 per level (max. 5). They regenerate at the start of each battle. | 4 levels |
| HEALING SPEED | -10 % time between heals per level. | 5 levels |
| RANGE | +5 % range per level. | 5 levels |
| FOCUS | +6 % damage, attack speed and mech speed per level to towers and mechs within its range. | 5 levels |
| ABDUCTOR: SECOND FARM | Buys a SECOND FARM below the first, at large size: twice the animals. When the first is full, captured animals go to the second. | 1 levels |
| CIRCLE OF GREED: MULTIPLIER | Monsters that die inside the circle of greed drop x2 loot and cards; each level raises the multiplier (x3, x4, x5). | 3 levels |
| CIRCLE OF GREED: AREA | +10 % circle of greed radius per level. | 5 levels |
| CIRCLE OF GREED: DURATION | +20 % time before the circle of greed disappears per level. | 5 levels |
